From the homely scuttle of coal at
the side of the hearth to the gorgeously verdant vegetation
of a forest of mammoth trees, might have appeared
a somewhat far cry in the eyes of those who lived some
fifty years ago. But there are few now who do
not know what was the origin of the coal which they
use so freely, and which in obedience to their demand
has been brought up more than a thousand feet from
the bowels of the earth; and, although familiarity
has in a sense bred contempt for that which a few
shillings will always purchase, in all probability
a stray thought does occasionally cross one’s
mind, giving birth to feelings of a more or less thankful
nature that such a store of heat and light was long
ago laid up in this earth of ours for our use, when
as yet man was not destined to put in an appearance
for many, many ages to come. We can scarcely
imagine the industrial condition of our country in
the absence of so fortunate a supply of coal; and
the many good things which are obtained from it, and
the uses to which, as we shall see, it can be put,
do indeed demand recognition.
Were our present forests uprooted
and overthrown, to be covered by sedimentary deposits
such as those which cover our coal-seams, the amount
of coal which would be thereby formed for use in some
future age, would amount to a thickness of perhaps
two or three inches at most, and yet, in one coal-field
alone, that of Westphalia, the 117 most important seams,
if placed one above the other in immediate succession,
would amount to no less than 294 feet of coal.
From this it is possible to form a faint idea of the
enormous growths of vegetation required to form some
of our representative coal beds. But the coal
is not found in one continuous bed. These numerous
seams of coal are interspersed between many thousands
of feet of sedimentary deposits, the whole of which
form the “coal-measures.” Now, each
of these seams represents the growth of a forest,
and to explain the whole series it is necessary to
suppose that between each deposit the land became
overwhelmed by the waters of the sea or lake, and
after a long subaqueous period, was again raised into
dry land, ready to become the birth-place of another
forest, which would again beget, under similarly repeated
conditions, another seam of coal. Of the conditions
necessary to bring these changes about we will speak
later on, but this instance is sufficient to show how
inadequate the quantity of fuel would be, were we
dependent entirely on our own existing forest growths.
However, we will leave for the present
the fascinating pursuit of theorising as to the how
and wherefore of these vast beds of coal, relegating
the geological part of the study of the carboniferous
system to a future chapter, where will be found some
more detailed account of the position of the coal-seams
in the strata which contain them. At present
the actual details of the coal itself will demand our
attention.
Coal is the mineral which has resulted,
after the lapse of thousands of thousands of years,
from the accumulations of vegetable material, caused
by the steady yearly shedding of leaves, fronds and
spores, from forests which existed in an early age;
these accumulated where the trees grew that bore them,
and formed in the first place, perhaps, beds of peat;
the beds have since been subjected to an ever-increasing
pressure of accumulating strata above them, compressing
the sheddings of a whole forest into a thickness in
some cases of a few inches of coal, and have been
acted upon by the internal heat of the earth, which
has caused them to part, to a varying degree, with
some of their component gases. If we reason from
analogy, we are compelled to admit that the origin
of coal is due to the accumulation of vegetation,
of which more scattered, but more distinct, representative
specimens occur in the shales and clays above and
below the coal-seams. But we are also able to
examine the texture itself of the various coals by
submitting extremely thin slices to a strong light
under the microscope, and are thus enabled to decide
whether the particular coal we are examining is formed
of conifers, horse-tails, club-mosses, or ferns, or
whether it consists simply of the accumulated sheddings
of all, or perhaps, as in some instances, of innumerable
spores.
In this way the structure of coal
can be accurately determined. Were we artificially
to prepare a mass of vegetable substance, and covering
it up entirely, subject it to great pressure, so that
but little of the volatile gases which would be formed
could escape, we might in the course of time produce
something approaching coal, but whether we obtained
lignite, jet, common bituminous coal, or anthracite,
would depend upon the possibilities of escape for
the gases contained in the mass.
Everybody has doubtless noticed that,
when a stagnant pool which contains a good deal of
decaying vegetation is stirred, bubbles of gas rise
to the surface from the mud below. This gas is
known as marsh-gas, or light carburetted hydrogen,
and gives rise to the ignis fatuus which hovers
about marshy land, and which is said to lure the weary
traveller to his doom. The vegetable mud is here
undergoing rapid decomposition, as there is nothing
to stay its progress, and no superposed load of strata
confining its resulting products within itself.
The gases therefore escape, and the breaking-up of
the tissues of the vegetation goes on rapidly.
The chemical changes which have taken
place in the beds of vegetation of the carboniferous
epoch, and which have transformed it into coal, are
even now but imperfectly understood. All we know
is that, under certain circumstances, one kind of
coal is formed, whilst under other conditions, other
kinds have resulted; whilst in some cases the processes
have resulted in the preparation of large quantities
of mineral oils, such as naphtha and petroleum.
Oils are also artificially produced from the so-called
waste-products of the gas-works, but in some parts
of the world the process of their manufacture has
gone on naturally, and a yearly increasing quantity
is being utilised. In England oil has been pumped
up from the carboniferous strata of Coalbrook Dale,
whilst in Sussex it has been found in smaller quantities,
where, in all probability, it has had its origin in
the lignitic beds of the Wealden strata. Immense
quantities are used for fuel by the Russian steamers
on the Caspian Sea, the Baku petroleum wells being
a most valuable possession. In Sicily, Persia,
and, far more important, in the United States, mineral
oils are found in great quantity.
In all probability coniferous trees,
similar to the living firs, pines, larches, &c., gave
rise for the most part to the mineral oils. The
class of living coniferae is well known for
the various oils which it furnishes naturally, and
for others which its representatives yield on being
subjected to distillation. The gradually increasing
amount of heat which we meet the deeper we go beneath
the surface, has been the cause of a slow and continuous
distillation, whilst the oil so distilled has found
its way to the surface in the shape of mineral-oil
springs, or has accumulated in troughs in the strata,
ready for use, to be drawn up when a well has been
sunk into it.
The plants which have gone to make
up the coal are not at once apparent to the naked
eye. We have to search among the shales and clays
and sandstones which enclose the coal-seams, and in
these we find petrified specimens which enable us
to build up in our mind pictures of the vegetable
creation which formed the jungles and forests of these
immensely remote ages, and which, densely packed together
on the old forest floor of those days, is now apparent
to us as coal.
A very large proportion of the plants
which have been found in the coal-bearing strata consists
of numerous species of ferns, the number of actual
species which have been preserved for us in our English
coal, being double the number now existing in Europe.
The greater part of these do not seem to have been
very much larger than our own living ferns, and, indeed,
many of them bear a close resemblance to some of our
own living species. The impressions they have
left on the shales of the coal-measures are most striking,
and point to a time when the sandy clay which imbedded
them was borne by water in a very tranquil manner,
to be deposited where the ferns had grown, enveloping
them gradually, and consolidating them into their
mass of future shale. In one species known as
the neuropteris, the nerves of the leaves are
as clear and as apparent as in a newly-grown fern,
the name being derived from two Greek words meaning
“nerve-fern.” It is interesting to
consider the history of such a leaf, throughout the
ages that have elapsed since it was part of a living
fern. First it grew up as a new frond, then gradually
unfolded itself, and developed into the perfect fern.
Then it became cut off by the rising waters, and buried
beneath an accumulation of sediment, and while momentous
changes have gone on in connection with the surface
of the earth, it has lain dormant in its hiding-place
exactly as we see it, until now excavated, with its
contemporaneous vegetation, to form fuel for our winter
fires.
Although many of the ferns greatly
resembled existing species, yet there were others
in these ancient days utterly unlike anything indigenous
to England now. There were undoubted tree-ferns,
similar to those which thrive now so luxuriously in
the tropics, and which throw out their graceful crowns
of ferns at the head of a naked stem, whilst on the
bark are the marks at different levels of the points
of attachment of former leaves. These have left
in their places cicatrices or scars, showing the
places from which they formerly grew. Amongst
the tree-ferns found are megaphyton, paloeopteris,
and caulopteris, all of which have these marks
upon them, thus proving that at one time even tree-ferns
had a habitat in England.
One form of tree-fern is known by
the name of Psaronius, and this was peculiar
in the possession of masses of aerial roots grouped
round the stem. Some of the smaller species exhibit
forms of leaves which are utterly unknown in the nomenclature
of living ferns. Most have had names assigned
to them in accordance with certain characteristics
which they possess. This was the more possible
since the fossilised impressions had been retained
in so distinct a manner. Here before us is a specimen
in a shale of pecopteris, as it is called,
(pekos, a comb). The leaf in some species
is not altogether unlike the well-known living fern
osmunda. The position of the pinnules
on both sides of the central stalk are seen in the
fossil to be shaped something like a comb, or a saw,
whilst up the centre of each pinnule the vein is as
prominent and noticeable as if the fern were but yesterday
waving gracefully in the air, and but to-day imbedded
in its shaly bed.
Sphenopteris, or “wedge-fern,”
is the name applied to another coal-fern; glossopteris,
or “tongue-leaf”; cyclopteris, or
“round-leaf”; odonlopteris, or “tooth-leaf,”
and many others, show their chief characteristics
in the names which they individually bear. Alethopteris
appears to have been the common brake of the coal-period,
and in some respects resembles pecopteris.
In some species of ferns so exact
are the representations which they have impressed
on the shale which contains them, that not only are
the veins and nerves distinctly visible, but even
the fructification still remains in the shape of the
marks left by the so-called seeds on the backs of the
leaves. Something more than a passing look at
the coal specimens in a good museum will well repay
the time so spent.
What are known as septarian nodules,
or snake-stones, are, at certain places, common in
the carboniferous strata. They are composed of
layers of ironstone and sandstone which have segregated
around some central object, such as a fern-leaf or
a shell. When the leaf of a fern has been found
to be the central object, it has been noticed that
the leaf can sometimes be separated from the stone
in the form of a carbonaceous film.
Experiments were made many years ago
by M. Goppert to illustrate the process of fossilisation
of ferns. Having placed some living ferns in a
mass of clay and dried them, he exposed them to a red
heat, and obtained thereby striking resemblances to
fossil plants. According to the degree of heat
to which they were subjected, the plants were found
to be either brown, a shining black, or entirely lost.
In the last mentioned case, only the impression remained,
but the carbonaceous matter had gone to stain the
surrounding clay black, thus indicating that the dark
colour of the coal-shales is due to the carbon derived
from the plants which they included.
Another very prominent member of the
vegetation of the coal period, was that order of plants
known as the Calamites. The generic distinctions
between fossil and living ferns were so slight in many
cases as to be almost indistinguishable. This
resemblance between the ancient and the modern is
not found so apparent in other plants. The Calamites
of the coal-measures bore indeed a very striking resemblance,
and were closely related, to our modern horse-tails,
as the equiseta are popularly called; but in
some respects they differed considerably.
Most people are acquainted with the
horse-tail (equisetum fluviatile) of our marshes
and ditches. It is a somewhat graceful plant,
and stands erect with a jointed stem. The foliage
is arranged in whorls around the joints, and, unlike
its fossil representatives, its joints are protected
by striated sheaths. The stem of the largest living
species rarely exceeds half-an-inch in diameter, whilst
that of the calamite attained a thickness of five
inches. But the great point which is noticeable
in the fossil calamites and equisetites
is that they grew to a far greater height than any
similar plant now living, sometimes being as much as
eight feet high. In the nature of their stems,
too, they exhibited a more highly organised arrangement
than their living representatives, having, according
to Dr Williamson, a “fistular pith, an exogenous
woody stem, and a thick smooth bark.” The
bark having almost al ways disappeared has left
the fluted stem known to us as the calamite. The
foliage consisted of whorls of long narrow leaves,
which differed only from the fern asterophyllites
in the fact that they were single-nerved. Sir
William Dawson assigns the calamites to four
sub-types: calamite proper, calamopitus,
calamodendron, and eucalamodendron.
Having used the word “exogenous,”
it might be as well to pay a little attention, in
passing, to the nomenclature and broad classification
of the various kinds of plants. We shall then
doubtless find it far easier thoroughly to understand
the position in the scale of organisation to which
the coal plants are referable.
The plants which are lowest in organisation
are known as Cellular. They are almost
entirely composed of numerous cells built up one above
the other, and possess none of the higher forms of
tissue and organisation which are met with elsewhere.
This division includes the lichens, sea-weeds, confervae
(green aquatic scum), fungi (mushrooms, dry-rot),
&c.
The division of Vascular plants
includes the far larger proportion of vegetation,
both living and fossil, and these plants are built
up of vessels and tissues of various shapes and character.
All plants are divided into (1) Cryptogams,
or Flowerless, such as mosses, ferns, equisetums,
and (2) Phanerogams, or Flowering. Flowering
plants are again divided into those with naked seeds,
as the conifers and cycads (gymnosperms), and those
whose seeds are enclosed in vessels, or ovaries (angiosperms).
Angiosperms are again divided into
the monocotyledons, as the palms, and dicotyledons,
which include most European trees.
Thus:
-------------------------------------------------------------------
| (M.A. Brongniart). | |(Lindley). |
|CELLULAR | | |
| Cryptogams (Flowerless) |Fungi, seaweeds, |Thallogens |
| |lichens | |
| | | |
|VASCULAR | | |
| Cryptogams (Flowerless) |Ferns, equisetums, |Acrogens |
| |mosses, lycopodiums | |
|PHANEROGAMS (Flowering) | | |
| Gymnosperms (having |Conifers and |Gymnogens |
| naked seeds) |cycads | |
| Two or more Cotyledons | | |
| Angiosperms (having | | |
| enclosed seeds) | | |
| Monocotyledons |Palms, lilies, |Endogens |
| |grasses | |
| Dicotyledons |Most European |Exogens |
| |trees and shrubs | |
-------------------------------------------------------------------
Adolphe Brongniart termed the coal
era the “Age of Acrogens,” because, as
we shall see, of the great predominance in those times
of vascular cryptogamic plants, known in Dr Lindley’s
nomenclature as “Acrogens.”
Two of these families have already
been dealt with, viz., the ferns (felices),
and the equisetums, (calamites and equisetites),
and we now have to pass on to another family.
This is that which includes the fossil representatives
of the Lycopodiums, or Club-mosses, and which goes
to make up in some coals as much as two-thirds of the
whole mass. Everyone is more or less familiar
with some of the living Lycopodiums, those delicate
little fern-like mosses which are to be found in many
a home. They are but lowly members of our British
flora, and it may seem somewhat astounding at first
sight that their remote ancestors occupied so important
a position in the forests of the ancient period of
which we are speaking. Some two hundred living
species are known, most of them being confined to
tropical climates. They are as a rule, low creeping
plants, although some few stand erect. There is
room for astonishment when we consider the fact that
the fossil representatives of the family, known as
Lepidodendra, attained a height of no less than
fifty feet, and, there is good ground for believing,
in many cases, a far greater magnitude. They
consist of long straight stems, or trunks which branch
considerably near the top. These stems are covered
with scars or scales, which have been caused by the
separation of the petioles or leaf-stalks, and
this gives rise to the name which the genus bears.
The scars are arranged in a spiral manner the whole
of the way up the stem, and the stems often remain
perfectly upright in the coal-mines, and reach into
the strata which have accumulated above the coal-seam.
Count Sternberg remarked that we are
unacquainted with any existing species of plant, which
like the Lepidodendron, preserves at all ages,
and throughout the whole extent of the trunk, the scars
formed by the attachment of the petioles, or
leaf-stalks, or the markings of the leaves themselves.
The yucca, dracaena, and palm, entirely shed their
scales when they are dried up, and there only remain
circles, or rings, arranged round the trunk in different
directions. The flabelliform palms preserve their
scales at the inferior extremity of the trunk only,
but lose them as they increase in age; and the stem
is entirely bare, from the middle to the superior
extremity. In the ancient Lepidodendron,
on the other hand, the more ancient the scale of the
leaf-stalk, the more apparent it still remains.
Portions of stems have been discovered which contain
leaf-scars far larger than those referred to above,
and we deduce from these fragments the fact that those
individuals which have been found whole, are not by
any means the largest of those which went to form so
large a proportion of the ancient coal-forests.
The lepidodendra bore linear one-nerved leaves,
and the stems always branched dichotomously and possessed
a central pith. Specimens variously named knorria,
lepidophloios, halonia, and ulodendron are
all referable to this family.
In some strata, as for instance that
of the Shropshire coalfield, quantities of elongated
cylindrical bodies known as lepidostrobi have
been found, which, it was early conjectured, were the
fruit of the giant club-mosses about which we have
just been speaking. Their appearance can be called
to mind by imagining the cylindrical fruit of the maize
or Indian corn to be reduced to some three or four
inches in length. The sporangia or cases which
contained the microscopic spores or seeds were arranged
around a central axis in a somewhat similar manner
to that in which maize is found. These bodies
have since been found actually situated at the end
of branches of lepidodendron, thus placing their
true nature beyond a doubt. The fossil seeds (spores)
do not appear to have exceeded in volume those of
recent club-mosses, and this although the actual trees
themselves grew to a size very many times greater than
the living species. This minuteness of the seed-germs
goes to explain the reason why, as Sir Charles Lyell
remarked, the same species of lepidodendra
are so widely distributed in the coal measures of Europe
and America, their spores being capable of an easy
transportation by the wind.
One striking feature in connection
with the fruit of the lepidodendron and other
ancient representatives of the club-moss tribe, is
that the bituminous coals in many, if not in most,
instances, are made up almost entirely of their spores
and spore-cases. Under a microscope, a piece of
such coal is seen to be thronged with the minute rounded
bodies of the spores interlacing one another and forming
almost the whole mass, whilst larger than these, and
often indeed enclosing them, are flattened bag-like
bodies which are none other than the compressed sporangia
which contained the former.
Now, the little Scottish or Alpine
club-moss which is so familiar, produces its own little
cones, each with its series of outside scales or leaves;
these are attached to the bags or spore-cases, which
are crowded with spores. Although in miniature,
yet it produces its fruit in just the same way, at
the terminations of its little branches, and the spores,
the actual germs of life, when examined microscopically,
are scarcely distinguishable from those which are
contained in certain bituminous coals. And, although
ancient club-mosses have been found in a fossilised
condition at least forty-nine feet high, the spores
are no larger than those of our miniature club-mosses
of the present day.
The spores are more or less composed
of pure bitumen, and the bituminous nature of the
coal depends largely on the presence or absence of
these microscopic bodies in it. The spores of
the living club-mosses contain so much resinous matter
that they are now largely used in the making of fireworks,
and upon the presence of this altered resinous matter
in coal depends its capability of providing a good
blazing coal.
At first sight it seems almost impossible
that such a minute cause should result in the formation
of huge masses of coal, such an inconceivable number
of spores being necessary to make even the smallest
fragment of coal. But if we look at the cloud
of spores that can be shaken from a single spike of
a club-moss, then imagine this to be repeated a thousand
times from each branch of a fairly tall tree, and then
finally picture a whole forest of such trees shedding
in due season their copious showers of spores to earth,
we shall perhaps be less amazed than we were at first
thought, at the stupendous result wrought out by so
minute an object.
Another well-known form of carboniferous
vegetation is that known as the Sigillaria,
and, connected with this form is one, which was long
familiar under the name of Stigmaria, but which
has since been satisfactorily proved to have formed
the branching root of the sigillaria. The older
geologists were in the habit of placing these plants
among the tree-ferns, principally on account of the
cicatrices which were left at the junctions of
the leaf-stalks with the stem, after the former had
fallen off. No foliage had, however, been met
with which was actually attached to the plants, and
hence, when it was discovered that some of them had
long attenuated leaves not at all like those possessed
by ferns, geologists were compelled to abandon this
classification of them, and even now no satisfactory
reference to existing orders of them has been made,
owing to their anomalous structure. The stems
are fluted from base to stem, although this is not
so apparent near the base, whilst the raised prominences
which now form the cicatrices, are arranged at
regular distances within the vertical grooves.
When they have remained standing for
some length of time, and the strata have been allowed
quietly to accumulate around the trunks, they have
escaped compression. They were evidently, to a
great extent, hollow like a reed, so that in those
trees which still remain vertical, the interior has
become filled up by a coat of sandstone, whilst the
bark has become transformed into an envelope of an
inch, or half an inch of coal. But many are found
lying in the strata in a horizontal plane. These
have been cast down and covered up by an ever-increasing
load of strata, so that the weight has, in the course
of time, compressed the tree into simply the thickness
of the double bark, that is, of the two opposite sides
of the envelope which covered it when living.
Sigillarae grew to a very great
height without branching, some specimens having measured
from 60 to 70 feet long. In accordance with their
outside markings, certain types are known as syringodendron,
favularia, and clathraria. Diploxylon
is a term applied to an interior stem referable to
this family.
But the most interesting point about
the sigillariae is the root. This was
for a long time regarded as an entirely distinct individual,
and the older geologists explained it in their writings
as a species of succulent aquatic plant, giving it
the name of stigmaria. They realized the
fact that it was almost universally found in those
beds which occur immediately beneath the coal seams,
but for a long time it did not strike them that it
might possibly be the root of a tree. In an old
edition of Lyell’s “Elements of Geology,”
utterly unlike existing editions in quality, quantity,
or comprehensiveness, after describing it as an extinct
species of water-plant, the author hazarded the conjecture
that it might ultimately be found to have a connection
with some other well-known plant or tree. It
was noticed that above the coal, in the roof, stigmariae
were absent, and that the stems of trees which occurred
there, had become flattened by the weight of the overlying
strata. The stigmariae on the other hand, abounded
in the underclay, as it is called, and were
not in any way compressed but retained what appeared
to be their natural shape and position. Hence
to explain their appearance, it was thought that they
were water-plants, ramifying the mud in every direction,
and finally becoming overwhelmed and covered by the
mud itself. On botanical grounds, Brongniart
and Lyell conjectured that they formed the roots of
other trees, and this became the more apparent as it
came to be acknowledged that the underclays were really
ancient soils. All doubt was, however, finally
dispelled by the discovery by Mr Binney, of a sigillaria
and a stigmaria in actual connection with each other,
in the Lancashire coal-field.
Stigmariae have since been found in
the Cape Breton coal-field, attached to Lepidodendra,
about which we have already spoken, and a similar
discovery has since been made in the British coal-fields.
This, therefore, would seem to shew the affinity of
the sigillaria to the lepidodendron, and through
it to the living lycopods, or club-mosses.
Some few species of stigmarian roots
had been discovered, and various specific names had
been given to them before their actual nature was made
out. What for some time were thought to be long
cylindrical leaves, have now been found to be simply
rootlets, and in specimens where these have been removed,
the surface of the stigmaria has been noticed to be
covered with large numbers of protuberant tubercles,
which have formed the bases of the rootlets.
There appears to have also been some special kind of
arrangement in their growth, since, unlike the roots
of most living plants, the tubercles to which these
rootlets were attached, were arranged spirally around
the main root. Each of these tubercles was pitted
in the centre, and into these the almost pointed ends
of the rootlets fitted, as by a ball and socket joint.
“A single trunk of sigillaria
in an erect forest presents an epitome of a coal-seam.
Its roots represent the stigmaria underclay;
its bark the compact coal; its woody axis, the mineral
charcoal; its fallen leaves and fruits, with remains
of herbaceous plants growing in its shade, mixed with
a little earthy matter, the layers of coarse coal.
The condition of the durable outer bark of erect trees,
concurs with the chemical theory of coal, in showing
the especial suitableness of this kind of tissue for
the production of the purer compact coals.” (Dawson,
“Structures in Coal.”)
There is yet one other family of plants
which must be mentioned, and which forms a very important
portion of the constituent flora of the coal
period. This is the great family of the coniferae,
which although differing in many respects from the
highly organised dicotyledons of the present day,
yet resembled them in some respects, especially in
the formation of an annual ring of woody growth.
The conifers are those trees which,
as the name would imply, bear their fruit in the form
of cones, such as the fir, larch, cedar, and others.
The order is one which is familiar to all, not only
on account of the cones they bear, and their sheddings,
which in the autumn strew the ground with a soft carpet
of long needle-like leaves, but also because of the
gum-like secretion of resin which is contained in their
tissues. Only a few species have been found in
the coal-beds, and these, on examination under the
microscope, have been discovered to be closely related
to the araucarian division of pines, rather than to
any of our common firs. The living species of
this tree is a native of Norfolk Island, in the Pacific,
and here it attains a height of 200 feet, with a girth
of 30 feet. From the peculiar arrangement of
the ducts in the elongated cellular tissue of the
tree, as seen under the microscope, the fossil conifers,
which exhibit this structure, have been placed in the
same division.
The familiar fossil known to geologists
as Sternbergia has now been shown to be the
cast of the central pith of these conifers, amongst
which may be mentioned cordaites, araucarites,
and dadoxylon.. The central cores had
become replaced with inorganic matter after the pith
had shrunk and left the space empty. This shrinkage
of the pith is a process which takes place in many
plants even when living, and instances will at once
occur, in which the stems of various species of shrubs
when broken open exhibit the remains of the shrunken
pith, in the shape of thin discs across the interval
cavity.
We might reasonably expect that where
we find the remains of fossil coniferous trees, we
should also meet with the cones or fruit which they
bear. And such is the case. In some coal-districts
fossil fruits, named cardiocarpum and trigonocarpum,
have been found in great quantities, and these have
now been decided by botanists to be the fruits of certain
conifers, allied, not to those which bear hard cones,
but to those which bear solitary fleshy fruits.
Sir Charles Lyell referred them to a Chinese genus
of the yew tribe called salisburia. Dawson
states that they are very similar to both taxus
and salisburia.. They are abundant in some
coal-measures, and are contained, not only in the coal
itself, but also in the sandstones and shales.
The under-clays appear to be devoid of them, and this
is, of course, exactly what might have been expected,
since the seeds would remain upon the soil until covered
up by vegetable matter, but would never form part
of the clay soil itself.
In connection with the varieties which
have been distinguished in the families of the conifers,
calamites, and sigillariae, Sir William Dawson
makes the following observations: “I believe
that there was a considerably wide range of organisation
in cordaitinae as well as in calamites
and sigillariae, and that it will eventually
be found that there were three lines of connection
between the higher cryptogams (flowerless) and the
phaenogams (flowering), one leading from the lycopodes
by the sigillariae, another leading by the cordaites,
and the third leading from the equisetums by
the calamites. Still further back the
characters, afterwards separated in the club-mosses,
mare’s-tails, and ferns, were united in the rhizocarps,
or, as some prefer to call them, the heterosporous
filicinae.”
In concluding this chapter dealing
with the various kinds of plants which have been discovered
as contributing to the formation of coal-measures,
it would be as well to say a word or two concerning
the climate which must have been necessary to permit
of the growth of such an abundance of vegetation.
It is at once admitted by all botanists that a moist,
humid, and warm atmosphere was necessary to account
for the existence of such an abundance of ferns.
The gorgeous waving tree-ferns which were doubtless
an important feature of the landscape, would have
required a moist heat such as does not now exist in
this country, although not necessarily a tropical
heat. The magnificent giant lycopodiums cast
into the shade all our living members of that class,
the largest of which perhaps are those that flourish
in New Zealand. In New Zealand, too, are found
many species of ferns, both those which are arborescent
and those which are of more humble stature. Add
to these the numerous conifers which are there found,
and we shall find that a forest in that country may
represent to a certain extent the appearance presented
by a forest of carboniferous vegetation. The ferns,
lycopods, and pines, however, which appear there,
it is but fair to add, are mixed with other types
allied to more recent forms of vegetation.
There are many reasons for believing
that the amount of carbonic acid gas then existing
in the atmosphere was larger than the quantity which
we now find, and Professor Tyndall has shown that
the effect of this would be to prevent radiation of
heat from the earth. The resulting forms of vegetation
would be such as would be comparable with those which
are now reared in the green-house or conservatory
in these latitudes. The gas would, in fact, act
as a glass roof, extending over the whole world.